WORKLIST ENTRIES (1):

DEUTEROLYSIN View alignment View Structure    Deuterolysin metalloprotease (M35) family signature
 Type of fingerprint: COMPOUND with 5  elements
Links:
   PRINTS; PR00756 ALADIPTASE; PR00791 PEPDIPTASEA; PR00730 THERMOLYSIN
   PRINTS; PR00787 NEUTRALPTASE; PR00782 LSHMANOLYSIN; PR00931 MICOLLPTASE
   PRINTS; PR00997 FRAGILYSIN; PR00786 NEPRILYSIN; PR00765 CRBOXYPTASEA
   PRINTS; PR00932 AMINO1PTASE; PR00789 OSIALOPTASE; PR00933 BLYTICPTASE
   PRINTS; PR00934 XHISDIPTASE; PR00919 THERMOPTASE; PR00998 CRBOXYPTASET   
   PRINTS; PR00999 FUNGALYSIN; PR01000 SREBPS2PTASE
   INTERPRO; IPR001384
   PROSITE; PS00142 ZINC_PROTEASE

 Creation date 30-JAN-1997; UPDATE 07-JUN-1999

   1. RAWLINGS, N.D. AND BARRETT, A.J.
   Evolutionary families of metallopeptidases.
   METHODS ENZYMOL. 248 183-228 (1995).

   2. TATSUMI, H., MURAKAMI, S., TSUJI, R.F., ISHIDA, Y., MURAKAMI, K., MASAKI,
   A., KAWABE, H., ARIMURA., H., NAKANO, E. AND MOTAI, H.
   Cloning and expression in yeast of a cDNA clone encoding Aspergillus oryzae
   neutral protease II, a unique metalloprotease.
   MOL.GEN.GENET. 228 97-103 (1991).

   3. MATSUMOTO, K., YAMAGUCHI, M. AND ICHISHIMA, E.
   Molecular cloning and nucletide sequence of the complementary DNA for
   penicillinolysin gene plnC, and 18kDa metalloendopeptidase gene from 
   Penicillium citrinum.
   BIOCHIM.BIOPHYS.ACTA 1218(3) 469-472 (1994).

   Metalloproteases are the most diverse of the four main types of protease,
   with more than 30 families identified to date [1]. Of these, around
   half contain the HEXXH motif, which has been shown in crystallographic
   studies to form part of the metal-binding site [1]. This HEXXH motif is 
   relatively common, but can be more stringently defined for metallo-
   proteases as abXHEbbHbc, where a is most often valine or threonine and 
   forms part of the S1' subsite in thermolysin and neprilysin, b is an
   uncharged residue, and c a hydrophobic residue. Proline is never found
   in this site, possibly because it would break the helical structure
   adopted by this motif in metalloproteases [1].
   
   Metalloproteases can be split into five groups on the basis of their metal-
   binding residues: the first three contain the HEXXH motif, the other two
   do not [1]. In the first group, a glutamic acid completes the active site -
   these are termed HEXXH+E: all families in this group show some sequence
   relationship and have been assigned to clan MA [1]. The second group, which
   have a third histidine as the extra metal-binding residue, are termed
   HEXXH+H and are grouped into clan MB on the basis of their inter-relation-
   ship [1]. In the third group, the additional metal-binding residues are
   unidentified. The fourth group is diverse - the metal-binding residues are
   known but do not form the HEXXH motif. And the fifth group comprises the
   remaining families where the metal-binding residues are as yet unknown [1].
   
   Deuterolysin is a microbial zinc-containing metalloprotease that shows 
   some similarity to thermolysin [2]. The protein is expressed with a
   possible 19-residue signal sequence, a 155-residue propeptide, and an
   active peptide of 177 residues [3]. The latter contains an HEXXH motif
   towards the C-terminus, but the other zinc ligands are as yet undetermined
   [2,3].
   
   DEUTEROLYSIN is a 5-element fingerprint that provides a signature for the
   deuterolysin (M35) family of metalloproteases. The fingerprint was derived
   from an initial alignment of 3 sequences: the motifs were drawn from
   virtually the full length of the active peptide, motif 1 containing a
   conserved Glu residue that may contribute to zinc binding, and motif 4
   including the region encoded by PROSITE pattern ZINC_PROTEASE (PS00142),
   which contains the HEXXH motif. Two iterations on OWL29.3 were required to
   reach convergence, at which point a true set comprising 4 sequences was
   identified.
  
   An update on SPTR37_9f identified a true set of 4 sequences.

  SUMMARY INFORMATION
      4 codes involving  5 elements
      0 codes involving  4 elements
      0 codes involving  3 elements
      0 codes involving  2 elements

   COMPOSITE FINGERPRINT INDEX
  
    5|   4    4    4    4    4  
    4|   0    0    0    0    0  
    3|   0    0    0    0    0  
    2|   0    0    0    0    0  
   --+--------------------------
     |   1    2    3    4    5  

True positives..
 ME24_ASPFL     PLNC_PENCI     NPII_ASPOR     Q09016         


  PROTEIN TITLES
   ME24_ASPFL       24 KD METALLOPROTEINASE PRECURSOR (EC 3.4.24.39) (DEUTEROLYS
   PLNC_PENCI       PENICILLOLYSIN PRECURSOR (EC 3.4.24.39) (DEUTEROLYSIN) - PEN
   NPII_ASPOR       NEUTRAL PROTEASE II PRECURSOR (EC 3.4.24.39) (DEUTEROLYSIN) 
   Q09016           MEP20 - ASPERGILLUS FUMIGATUS (SARTORYA FUMIGATA).

SCAN HISTORY OWL29_3 2 100 NSINGLE SPTR37_9f 2 5 NSINGLE INITIAL MOTIF SETS DEUTEROLYSIN1 Length of motif = 30 Motif number = 1 Deuterolysin metalloprotease motif I - 1 PCODE ST INT LSNTVKLANAAATAARSGSASKFSEYFKTT PLNC_PENCI 192 192 LKNTVSLANQAASAAQSGSSSRFQEYFKTT ME24_ASPFL 211 211 LSNAAKLANQAAEAAESGDESKFEEYFKTT NPII_ASPOR 193 193 DEUTEROLYSIN2 Length of motif = 26 Motif number = 2 Deuterolysin metalloprotease motif II - 1 PCODE ST INT RSVVAARLEAVAKEAQSASSGSTTYY PLNC_PENCI 226 4 RTSVAARFRAVASEASSTSSGSTTYY ME24_ASPFL 245 4 RTTVAERLRAVAKEAGSTSGGSTTYH NPII_ASPOR 227 4 DEUTEROLYSIN3 Length of motif = 30 Motif number = 3 Deuterolysin metalloprotease motif III - 1 PCODE ST INT CSDTLGYCETNVLAYTLPARNIIANCDIYY PLNC_PENCI 252 0 CTDTYGYCSSNVPGVHLPAYNIIANCDIYY ME24_ASPFL 271 0 CNDPYGYCEPNVLAYTLPSKNEIANCDIYY NPII_ASPOR 253 0 DEUTEROLYSIN4 Length of motif = 29 Motif number = 4 Deuterolysin metalloprotease motif IV - 1 PCODE ST INT LAGTCHQQDQATTTLHEFTHAPGVYSPGT PLNC_PENCI 287 5 LTRTCHAQDQATTTLHEFTHAPGVYSPGT ME24_ASPFL 306 5 LAQKCHAQDQATTTLHEFTHAPGVYQPGT NPII_ASPOR 288 5 DEUTEROLYSIN5 Length of motif = 30 Motif number = 5 Deuterolysin metalloprotease motif V - 1 PCODE ST INT DDLGYGYSAATSLSSSQAVLNADSYALYAN PLNC_PENCI 316 0 DDLGYGYDAATALSSSQALNNVDTYALFAN ME24_ASPFL 335 0 EDLGYGYDAATQLSAQDALNNADSYALYAN NPII_ASPOR 317 0 FINAL MOTIF SETS DEUTEROLYSIN1 Length of motif = 30 Motif number = 1 Deuterolysin metalloprotease motif I - 2 PCODE ST INT LKNTVSLANQAASAAQSGSSSRFQEYFKTT ME24_ASPFL 211 211 LSNTVKLANAAATAARSGSASKFSEYFKTT PLNC_PENCI 192 192 LSNAAKLANQAAEAAESGDESKFEEYFKTT NPII_ASPOR 193 193 LRNAGSLANAAASAASSGSSTRFQEYFKTT Q09016 190 190 DEUTEROLYSIN2 Length of motif = 26 Motif number = 2 Deuterolysin metalloprotease motif II - 2 PCODE ST INT RTSVAARFRAVASEASSTSSGSTTYY ME24_ASPFL 245 4 RSVVAARLEAVAKEAQSASSGSTTYY PLNC_PENCI 226 4 RTTVAERLRAVAKEAGSTSGGSTTYH NPII_ASPOR 227 4 PENVGGRFRAVGREASSQSSGKTTYY Q09016 223 3 DEUTEROLYSIN3 Length of motif = 30 Motif number = 3 Deuterolysin metalloprotease motif III - 2 PCODE ST INT CTDTYGYCSSNVPGVHLPAYNIIANCDIYY ME24_ASPFL 271 0 CSDTLGYCETNVLAYTLPARNIIANCDIYY PLNC_PENCI 252 0 CNDPYGYCEPNVLAYTLPSKNEIANCDIYY NPII_ASPOR 253 0 CNDPYGYCDSNTLAYTLPSSNLIANCDIYY Q09016 249 0 DEUTEROLYSIN4 Length of motif = 29 Motif number = 4 Deuterolysin metalloprotease motif IV - 2 PCODE ST INT LTRTCHAQDQATTTLHEFTHAPGVYSPGT ME24_ASPFL 306 5 LAGTCHQQDQATTTLHEFTHAPGVYSPGT PLNC_PENCI 287 5 LAQKCHAQDQATTTLHEFTHAPGVYQPGT NPII_ASPOR 288 5 LTSSCHAQDQATTTLHEFTHAPAVYSPGT Q09016 284 5 DEUTEROLYSIN5 Length of motif = 30 Motif number = 5 Deuterolysin metalloprotease motif V - 2 PCODE ST INT DDLGYGYDAATALSSSQALNNVDTYALFAN ME24_ASPFL 335 0 DDLGYGYSAATSLSSSQAVLNADSYALYAN PLNC_PENCI 316 0 EDLGYGYDAATQLSAQDALNNADSYALYAN NPII_ASPOR 317 0 DDYAYGYRASTALSASQALLNADTYALFAN Q09016 313 0

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