WORKLIST ENTRIES (1):

GLHYDLASE14A View alignment View Structure    Bacterial beta-amylase signature
 Type of fingerprint: COMPOUND with 5  elements
Links:
   PRINTS; PR00131 GLHYDRLASE1; PR00132 GLHYDRLASE2; PR00133 GLHYDRLASE3
   PRINTS; PR00732 GLHYDRLASE4; PR00733 GLHYDRLASE6; PR00734 GLHYDRLASE7
   PRINTS; PR00735 GLHYDRLASE8; PR00134 GLHYDRLASE10; PR00911 GLHYDRLASE11
   PRINTS; PR00736 GLHYDRLASE15; PR00737 GLHYDRLASE16; PR00738 GLHYDRLASE20
   PRINTS; PR00739 GLHYDRLASE26; PR00740 GLHYDRLASE27; PR00741 GLHYDRLASE29
   PRINTS; PR00843 GLHYDRLASE30; PR00742 GLHYDRLASE35; PR00743 GLHYDRLASE36
   PRINTS; PR00744 GLHYDRLASE37; PR00745 GLHYDRLASE39; PR00746 GLHYDRLASE41
   PRINTS; PR00747 GLHYDRLASE47; PR00844 GLHYDRLASE48; PR00845 GLHYDRLASE52
   PRINTS; PR00846 GLHYDRLASE56; PR00849 GLHYDRLASE58; PR00850 GLHYDRLASE59
   PRINTS; PR00748 MELIBIASE; PR00137 LYSOZYME; PR00684 T4LYSOZYME
   PRINTS; PR00749 LYSOZYMEG; PR00110 ALPHAAMYLASE; PR00750 BETAAMYLASE
   PRINTS; PR00842 GLHYDLASE14B; PR00847 HYALURONDASE; PR00848 SPERMPH20
   INTERPRO; IPR000125
   PDB; 1BTC 3Dinfo
   SCOP; 1BTC
   CATH; 1BTC

 Creation date 11-MAR-1998; UPDATE 07-JUN-1999

   1. HENRISSAT, B.
   A classification of glycosyl hydrolases based on amino acid sequence
   similarities.
   BIOCHEM.J. 280 309-316 (1991).

   2. HENRISSAT, B. AND BAIROCH, A.
   New families in the classification of glycosyl hydrolases based on amino
   acid sequence similarities.
   BIOCHEM.J. 293 781-788 (1993).

   3. HENRISSAT, B. AND BAIROCH, A.
   Updating the sequence-based classification of glycosyl hydrolases.
   BIOCHEM.J. 316 695-696 (1996).

   4. EL HASSOUNI, M., HENRISSAT, B., CHIPPAUX, M. AND BARRAS, F.
   Nucleotide sequences of the Arb genes, which control beta-glucosidase
   utilisation in Erwinia chrysanthemi - Comparison with the Escherichia
   coli Bgl operon and evidence for a new beta-glycohydrolase family
   including enzymes from eubacteria, archaebacteria and humans.
   J.BACTERIOL. 174 765-777 (1992).
  
   5. VIKSONIELSEN, A., CHRISTENSEN, T.M., BOJKO, M. AND MARCUSSEN, J.
   Purification and characterization of beta-amylase from leaves of potato 
   (Solanum tuberosum).
   PHYSIOLOGIA PLANTARUM 99 190-196 (1997).

   6. MIKAMI, B., SATO, M., SHIBATA, T., HIROSE, M., AIBARA, S., KATSUBE, Y.
   AND MORITA, Y.
   Three-dimensional structure of soybean beta-amylase determined at 3.0 A 
   resolution: preliminary chain tracing of the complex with alpha-cyclodextrin. 
   J.BIOCHEM. 112 541-546 (1992). 

   O-Glycosyl hydrolases (EC 3.2.1.-) are a widespread group of enzymes that
   hydrolyse the glycosidic bond between two or more carbohydrates, or between
   a carbohydrate and a non-carbohydrate moiety. A classification system for
   glycosyl hydrolases, based on sequence similarity, has led to the definition
   of up to 60 different families [1-4] (http://expasy.hcuge.ch/cgi-bin/lists?
   glycosid.txt). Family 14 (EC 3.2.1.2) encompasses the beta-amylases.
  
   Beta-amylases, which are found in plants and bacteria, hydrolyse 1,4-alpha-
   glucosidic linkages in starch-type polysaccharide substrates, removing
   successive maltose units from the non-reducing ends of the chains [5]. In 
   potato plants, the enzyme has been found to work optimally at 40 degrees C, 
   becoming unstable above this temperature [5]. On the basis of sequence
   comparisons, plant and bacterial beta-amylases can be readily distinguished
   from each other. 
  
   The 3D structure of a complex of soybean beta-amylase with an inhibitor 
   (alpha-cyclodextrin) has been determined to 3.0A resolution by X-ray
   diffraction [6]. The enzyme folds into large and small domains: the large
   domain has a (beta alpha)8 super-secondary structural core, while the smaller
   is formed from two long loops extending from the beta-3 and beta-4 strands
   of the (beta alpha)8 fold [6]. The interface of the two domains, together
   with shorter loops from the (beta alpha)8 core, form a deep cleft, in which 
   the inhibitor binds [6]. Two maltose molecules also bind in the cleft,
   one sharing a binding site with alpha-cyclodextrin, and the other sitting
   more deeply in the cleft [6]. Sequence alignments allow us to relate
   features of the bacterial beta-amylases to the soybean structure.
   
   GLHYDLASE14A is a 5-element fingerprint that provides a signature for 
   the bacterial beta-amylases. The fingerprint was derived from an initial
   alignment of 4 sequences: the motifs were drawn from conserved sections
   spanning virtually the full alignment length, focusing on those regions
   that characterise the bacterial beta-amylases but distinguish them from
   the rest of the beta-amylase family - motif 1 spans helices 5 and 6, beta-
   strand 6 and the N-terminus of strand 7; motif 2 includes the C-terminal
   half of helix 12; and motif 3 includes helix 13. Two iterations on OWL30.0
   were required to reach convergence, at which point a true set comprising 6
   sequences was identified. A single partial match was also found, AB000264,
   a fragment from Bacillus firmus that lacks the portion of sequence bearing
   motif 5.
  
   An update on SPTR37_9f identified a true set of 4 sequences.

  SUMMARY INFORMATION
      4 codes involving  5 elements
      0 codes involving  4 elements
      0 codes involving  3 elements
      0 codes involving  2 elements

   COMPOSITE FINGERPRINT INDEX
  
    5|   4    4    4    4    4  
    4|   0    0    0    0    0  
    3|   0    0    0    0    0  
    2|   0    0    0    0    0  
   --+--------------------------
     |   1    2    3    4    5  

True positives..
 AMYB_BACCI     AMYB_PAEPO     AMYB_THETU     AMYB_BACCE     


  PROTEIN TITLES
   AMYB_BACCI       BETA-AMYLASE PRECURSOR (EC 3.2.1.2) (1,4-ALPHA-D-GLUCAN MALT
   AMYB_PAEPO       BETA/ALPHA-AMYLASE PRECURSOR [INCLUDES: BETA-AMYLASE (EC 3.2
   AMYB_THETU       BETA-AMYLASE, THERMOPHILIC PRECURSOR (EC 3.2.1.2) (1,4-ALPHA
   AMYB_BACCE       BETA-AMYLASE PRECURSOR (EC 3.2.1.2) (1,4-ALPHA-D-GLUCAN MALT

SCAN HISTORY OWL30_0 2 75 NSINGLE SPTR37_9f 2 6 NSINGLE INITIAL MOTIF SETS GLHYDLASE14A1 Length of motif = 20 Motif number = 1 Bacterial beta-amylase motif I - 1 PCODE ST INT WSKGSADEMQFKDESGYVNN AMYB_BACCI 136 136 SSKGSADEMQFKDESGYANS AMYB_BACPO 135 135 WTKDTQDNMQYKDEAGNWDN AMYB_THETU 132 132 WNQKSDDSLYFKSETGTVNK AMYB_BACCE 138 138 GLHYDLASE14A2 Length of motif = 16 Motif number = 2 Bacterial beta-amylase motif II - 1 PCODE ST INT TAMTTKYGSLDKINAA AMYB_BACCI 232 76 TAMNDKYGSLDKINAA AMYB_BACPO 231 76 NAMKSKYGTIAAVNSA AMYB_THETU 228 76 LWVLNKYGSLNEVNKA AMYB_BACCE 235 77 GLHYDLASE14A3 Length of motif = 18 Motif number = 3 Bacterial beta-amylase motif III - 1 PCODE ST INT WGTNLTSMSQISPPTDSD AMYB_BACCI 248 0 WGTKLTSLSQINPPTDGD AMYB_BACPO 247 0 WGTSLTDFSQISPPTDGD AMYB_THETU 244 0 WGTKLISELAILPPSDGE AMYB_BACCE 251 0 GLHYDLASE14A4 Length of motif = 22 Motif number = 4 Bacterial beta-amylase motif IV - 1 PCODE ST INT TRFGYNGFTLLRINNIVNSDGS AMYB_BACCI 413 147 TKFGYHGFTLLRINNLVNNDGS AMYB_BACPO 412 147 TGYNFSGFTLLRLSNIVNSDGS AMYB_THETU 412 150 FNYNFSGFTLLRYQDVMYNNSL AMYB_BACCE 416 147 GLHYDLASE14A5 Length of motif = 14 Motif number = 5 Bacterial beta-amylase motif V - 1 PCODE ST INT PGVKMPDSEISGYA AMYB_BACCI 490 55 PGVKMQDAEISGYA AMYB_BACPO 487 53 GPASCPNYPTWTIT AMYB_THETU 490 56 DSHSNDWRGNVVLP AMYB_BACCE 489 51 FINAL MOTIF SETS GLHYDLASE14A1 Length of motif = 20 Motif number = 1 Bacterial beta-amylase motif I - 2 PCODE ST INT WSKGSADEMQFKDESGYVNN AMYB_BACCI 136 136 SSKGSADEMQFKDESGYANS AMYB_PAEPO 135 135 WTKDTQDNMQYKDEAGNWDN AMYB_THETU 132 132 WNQKSDDSLYFKSETGTVNK AMYB_BACCE 138 138 GLHYDLASE14A2 Length of motif = 16 Motif number = 2 Bacterial beta-amylase motif II - 2 PCODE ST INT TAMTTKYGSLDKINAA AMYB_BACCI 232 76 TAMNDKYGSLDKINAA AMYB_PAEPO 231 76 NAMKSKYGTIAAVNSA AMYB_THETU 228 76 LWVLNKYGSLNEVNKA AMYB_BACCE 235 77 GLHYDLASE14A3 Length of motif = 18 Motif number = 3 Bacterial beta-amylase motif III - 2 PCODE ST INT WGTNLTSMSQISPPTDSD AMYB_BACCI 248 0 WGTKLTSLSQINPPTDGD AMYB_PAEPO 247 0 WGTSLTDFSQISPPTDGD AMYB_THETU 244 0 WGTKLISELAILPPSDGE AMYB_BACCE 251 0 GLHYDLASE14A4 Length of motif = 22 Motif number = 4 Bacterial beta-amylase motif IV - 2 PCODE ST INT TRFGYNGFTLLRINNIVNSDGS AMYB_BACCI 413 147 TKFGYHGFTLLRINNLVNNDGS AMYB_PAEPO 412 147 TGYNFSGFTLLRLSNIVNSDGS AMYB_THETU 412 150 FNYNFSGFTLLRYQDVMYNNSL AMYB_BACCE 416 147 GLHYDLASE14A5 Length of motif = 14 Motif number = 5 Bacterial beta-amylase motif V - 2 PCODE ST INT PGVKMPDSEISGYA AMYB_BACCI 490 55 PGVKMQDAEISGYA AMYB_PAEPO 487 53 GPASCPNYPTWTIT AMYB_THETU 490 56 DSHSNDWRGNVVLP AMYB_BACCE 489 51

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